Publications are listed by author, alphabetically and made available where possible.

Papers that have been submitted, accepted or In Press are indicated. The references D#.#.# refer to the project deliverables

Papers on Physarum computing in arXiv can be found by following the link here

Adamatzky, Andrew “Physarum Machines: Computers from Slime Mould”, World Scientific Series on Nonlinear Science, Series A (2010): ISBN-13:

Adamatzky, Andrew “Route 20, Autobahn 7, and Slime Mold: Approximating the Longest Roads in USA and Germany With Slime Mold on 3-D Terrains”, IEEE Transactions Cybernetics. (2013) In Press – D1.2.1.

Adamatzky, Andrew. “Slime mould tactile sensor.” Sensors and Actuators B: Chemical Volume 188, November (2013): 38–44. – D.1.2.3.

Adamatzky, Andrew. “Tactile bristle sensors made with slime mould.” Sensors Journal, IEEE (2013) DOI: 10.1109/JSEN.2013.2283192 (previous version   PDF Tactile Bristle Sensors Made With Slime Mould D.1.2.4

Adamatzky, Andrew. “Physarum wires: Self-growing self-repairing smart wires made from slime mould”, Submitted (2013)

Adamatzky, Andrew. “Physarum oscillator.”(2013), Submitted

Adamatzky, Andrew “Slime mould electronic oscillators” Microelectronic Engineering Volume 124 25 July 2014 p58-65 DOI: 10.1016/j.mee.2014.04.022

Adamatzky, Andrew. “Towards slime mould colour sensor: Recognition of colours by Physarum polycephalum.” Organic Electronics. Volume 14, Issue 12, December (2013): 3355–3361

Adamatzky, Andrew, & R Armstrong., J Jones., G Yukio. “Creativity of slime mould.” International  Journal of General Systems (2013), Vol.42, Issue 5, p441-457- D9.1.1.

Adamatzky, Andrew,  & Victor Erokhin, Martin Grube, Theresa Schubert, Andrew Schumann. “Physarum Chip Project: Growing Computers From Slime Mould”, International Journal of Unconventional Computing, Vol. 8, No. 4 (2012): 319-323. Abstract PDF: Physarum Chip Project Growing Computers from Slime Mould

Adamatzky, A. and Jones, J. “On using compressibility to detect when slime mould completed computation”, Complexity. DOI:10.1002/cplx.21645. -D.6.1.5.

Adamatzky, Andrew, & G Martinez. “Bioimitation of Mexican migration routes to the USA with slime mould on 3D terrains” J Bionic Engineering 10   (2013): 242–250,  – D1.2.2.

Adamatzky, Andrew. & Richard Mayne “Actin automata: Phenomenology and localizations” 2014

Adamatzky, Andrew, & Richard Mayne. “Toward Hybrid Nanostructure-Slime Mould Devices” Nano Life 2014 DOI: 10.1142/S179398441450007X

Adamatzky, Andrew, & T Schubert “Slime mould microfluidic logical gates.” Materials Today  Volume 17 Issue 2, March 2014 p86-91  doi:10.1016/j.mattod.2014.01.018. D5.3.1.

Adamatzky, Andrew, & James Whiting, Ben De Lacy Costello. “Sensory fusion in Physarum polycephalum and implementing multi-sensory functional computation” Biosystems  Volume 119 May 2014 p45-52 DOI: 10.1016/j.biosystems.2014.03.003

Adamatzky, Andrew. & James Whiting, Ben De Lacy Costello. “Slime mould logic gates based on frequency changes of electrical potential oscillation” Biosystems Volume 124 October 2014 p21-25. DOI: 10.1016/j.biosystems.2014.08.001

Adamatzky, Andrew., & Xiaoge Zhang, Yajuan Zhang, Zili Zhang, Sankaran Mahadevan, Yong Deng. “Rapid Physarum Algorithm for the shortest path problem” Applied Soft Computing Volume 23, October 2014  DOI: 10.1016/j.asoc.2014.05.032

Berzina, Tatiana “Organic memristor device: fabrication, working principles and interfacing with living systems for unconventional computing.” Proc. E-MRS 2014 Spring meeting, Physica Status solidi C, (2014) Accepted – D8.1.4.

Berzina, T. & A. Dimonte, A. Cifarelli, and V. Erokhin. “Hybrid slime mold-based system for unconventional computing”, International Journal of General Systems, (2015). In press. -D8.3.2.

Bull, Larry, &  A Adamatzky. “Evolving gene regulatory networks with mobile DNA mechanisms” Computational Intelligence (UKCI), (2013)

Cifarelli, Angelica, & Tatiana Berzina, Victor Erokhin. “BIO-ORGANIC MEMRISTORS DEVICE: POLYANILINE-PHYSARUM POLYCEPHALUM INTERFACE.” Proc. E-MRS 2014 Spring meeting, Physica Status solidi C, (2014), Vol.12, p218-225. – D8.1.3.

Cifarelli, Angelica, &  Alice Dimonte, Tatiana Berzina, Victor Erokhin. “Non-liner bioelectronic element: Schottky effect and electrochemistry.” International Journal of Unconventional Computing, (2014), Vol.10, p375-379. – D.5.3.3.

Cifarelli, Angelica, & Alice Dimonte, Tatiana Berzina, Victor Erokhin. “On Loading of Slime Mould Physarum polycephalum with Microparticles for Unconventional Computing Application” BioNanoScience DOI: 10.1007/s12668-013-0124-3 (2014). – D8.1.2.

De Lacy Costello, Ben, & H. Al Kateb, N. Ratcliffe.  “Identifcation of the volatile secretions of the plasmodium of Physarum Polycephalum” (2013), In Press. – D1.1.1.

De Lacy Costello, Ben,& Andrew Adamatzky. “Assessing the chemotaxis behaviour of Physarum Polycephalum to a range of simple VOCs” (2013), Submitted – D1.3.1.

Dimonte, A & T. Berzina, A. Cifarelli, V. Chiesi, F. Albertini, and V. Erokhin. “Conductivity patterning with Physarum polycephalum: Natural growth and deflecting”, Physica Status Solidi C, Vol.12, p197-201. -D8.3.1.

Dimonte, Alice & Berzina, T., Pavesi, M., & Erokhin, V. “Hysteresis loop and cross-talk of organic memristive devices”, Microelectronics Journal, Vol.45, Issue11, p1396-1400. DOI:10.1016/j.mejo.2014.09.009. D8.1.1.

Dimonte, Alice & Tatiana Berzina, Victor Erokhin. “On the influence of music on Physarum polycephalum’s foreaging behaviour”. To be submitted. -D9.2.1.

Dimonte, Alice, & Angelica Cifarelli, Tatiana Berzina, V. Chiesi,P. Ferro, T. Besagni, F. Albertini, Andrew Adamatzky, Victor Erokhin. “Magnetic nanoparticles – loaded Physarum polycephalum: directed growth and particles distribution” Interdisciplinary Sciences- Computational Life Sciences, (2014) Submitted-  D5.1.1.

Dimonte, Alice, & Tatitana Berzina, Victor Erokhin. “Physarum computing: DEFLECTOR based on slime mold magnetic nanoparticle composite” Proc. E-MRS 2014 Spring meeting, Physica Status solidi C, (2014) Accepted – D.5.3.4.

Dimonte, A. & F. Fermi, T. Berzina, and V. Erokhin. “Spectral imaging method for studying Physarum polycephalum growth on polyaniline surface”, Materials Science and Engineering:C, (2015). Under reviewing. -D8.3.3.

Erokhin, Victor. “Organic Memristive Devices: Architecture, Properties and Applications in Neuromorphic Networks” Proceedings of the 2013 IEEE International Conference on Electronics, Circuits, and Systems,ICECS2013, December 8-11 2013, Abu Dhabi, UAE, Paper ID 6352 (2013)  – D8.1.1.

Erokhin, Victor. “Organic Memristive Devices and Neuromorphic circuits”, Memristor Networks, A.Adamatzky and L. Chua (Eds.), Springer, Cham, Heidelberg, New York, Dordrecht, London (2014).  – D8.1.2.

Gale E.,& Ben De Lacy Costello,Andrew Adamatzky. “Are Slime Moulds Living Memristors?” (2013), In Press –  D5.3.2.

Garcia, Leslie & Theresa Schubert.Sonification 1“. -D9.2.5.

Garcia, Leslie & Theresa Schubert.Sonification 2“. -D9.2.6.

Grube M. “A simple platform for interfacing plasmodial networks”. (2015). To be submitted. D8.2.4.

Jones, Jeff. “Mechanisms Inducing Parallel Computation in a Model of Physarum polycephalum Transport Networks”, Submitted. -D6.1.1.

Jones, Jeff. “A Morphological Adaptation Approach to Path Planning Inspired by Slime Mould”, International Journal of General Systems. DOI: 10.1080/03081079.2014.997526. -D7.1.5.

Jones, Jeff. “Towards Lateral Inhibition and Collective Perception in Unorganised Non-Neural Systems”, Computational Intelligence, Medecine and Biology: Selected Links. Submitted. -D6.1.7.

Jones, Jeff. “A Virtual Material Approach to Morphological Computation”,  Opinions and Outlooks on Morphological Computation, Eds: H. Hauser, R.M. Fuchslin and R. Pfeifer, University of Zurich. (2014) In Press. -D6.1.2

Jones, Jeff. “Material-based Non-neural Analogues of Lateral Inhibition: A Multi-agent Approach”, Proceedings of Symposium: Adaptive Materials, Devices and Systems Towards Unconventional Computing and Robotics: Modelling and Implementation, Rhodes, September 2014. -D6.1.6.

Jones, Jeff. “Exploiting Environmental Computation in a Multi-Agent Model of Slime Mould”, Proceedings of Symposium: 1st International Symposium on Artificial, Biological and Bio-Inspired Intelligence (ABBII)Rhodes, September 2014. -D6.1.3.

Jones, Jeff. “Sketching by Unconventional Computation: From Collective Perception to Collective Misperception”, AISB symposium on Music, Arts and Unconventional Computing, Canterbury UK. (Submitted). -D6.1.8.

Jones, Jeff. ““, (2014). -D9.2.2.2.

Jones, Jeff. “” (2014). -D9.2.3.

Jones, Jeff. “” (2015). -D9.2.4.

Jones, Jeff, &  Andrew Adamatzky. “Computation of the travelling salesman problem by a shrinking blob” Natural Computing, (13), 1, p. 1-16, (2014).

Jones, Jeff, & Andrew Adamatzky. “Slime Mould Inspired Generalised Voronoi Diagrams via Attraction and Repulsion Fields”, International  Journal of Bifurcation and Chaos (2013), In Press. – D2.1.1.

Jones, Jeff,  & Andrew Adamatzky. “Material Approximation of Data Smoothing and Spline Curves Inspired by Slime Mould”, Bioinspiration and Biomimetics, 9, (3), 036013, (2014) . -D7.1.1.

Jones, Jeff, & Andrew Adamatzky. “Approximation of Statistical Analysis and Estimation by Morphological Adaptation in a Model of Slime Mould”, International Journal of Unconventional Computing (2014)  In-press. -D7.1.2.

Jones, J. & Richard Mayne, Andrew Adamatzky. “Representation of Shape Mediated by Environmental Stimuli in Physarum polycephalum and a Multi-agent Model”. Submitted. -D7.1.4.

Mayne, Richard & Andrew Adamatzky. “Slime mould foraging behaviour as optically coupled logical operations”, International Journal of General Systems, Taylor and Francis, London, UK, (2015), p305-313. DOI: 10.1080/03081079.2014.997528. -D2.2.3

Mayne, Richard, & Andrew Adamatzky. “Towards Hybrid Artificial-Slime Mould Devices”, Biophysical Reviews and Letters  (2013) Submitted

Mayne, Richard, & Andrew Adamatzky. “Towards hybrid nanostructure-slime mould devices”, Nano LIFE, World Scientific, London, UK (2015). DOI:

Mayne, Richard & Andrew Adamatzky, Jeff Jones. “On the role of the plasmodial cytoskeleton in facilitating intelligent behaviour in slime mould Physarum polycephalum”. Communicative and Integrative Biology. Vol.7. DOI: -D.2.2.4

Mayne, Richard, & David Patton, Ben de Lacy Costello, Rosemary Camilla Patton, Andrew Adamatzky. “On loading slime mould Physarum polycephalum with metallic particles.” (2013) Submitted

Mayne, Richard, & David Patton, Ben de Lacy Costello, Andrew Adamatzky, Rosemary Camilla Patton “On  the internalisation, intraplasmodial carriage and excretion of metallic nanoparticles in the slime mould Physarum polycephalum.” International  Journal of Molecular and NanoComputing (2013)

Mayne, Richard, & Michail-Antisthenis Tsompanas, Georgios Ch. Sirakoulis, Andrew Adamatzky. “Towards a slime mould-FPGA interface”, Biomedical Engineering Letters, Springer, London, UK (2015), p51-57. DOI: – D.2.2.1

Pancerz, K., Schumann, A. “Rough Set Models of Physarum Machines”, International Journal of General Systems. DOI:10.1080/03081079.2014.997529. -D7.2.1

Pancerz, K., Schumann, A. “Some Issues on an Object-Oriented Programming Language for Physarum Machines”, Applications of Computational Intelligence in Biomedical Technology. Studies in Computational Intelligence, Springer International Publishing. Accepted. -D7.2.4

Pancerz, K., Schumann, A. “Principles of an Object-Oriented Programming Language for Physarum Polycephalum Computing”, Proceedings of the 10th International Conference on Digital Technologies (DT’2014), Zilina, Slovakia, July 9-11, 2014, pp. 273-280. -D7.2.5.

Phychip Interactive Online Publication. “Slime Mould Atlas“. -D9.4.1.

Romeo, A &, A. Dimonte, G. Tarabella, P. D’Angelo, V. Erokhin, and S. Iannotta. “A bio-inspired memory device based on interfacing Physarum polycephalum with an organic semiconductor”, APL Materials, Vol.3. -D8.2.2.

Schubert, Theresa. “Paper on interfacing PhyChips with humans via slime mould” (Chroma+Phy – a living wearable connecting humans and their environment, Living Machines) LNAI 8064,  Springer, Heidelberg (2013): 444-446  DOI 10.1007/978-3-642-39802-5_61 Chroma+Phy A living wearable connecting humans and their environment

Schubert, Theresa. Video of RGB sensitivity Growing Kolmogorov-Uspensky Machines (2014). -D9.1.2.

Schubert, Theresa. Video Growing Geometries II. -D9.1.3.

Schubert, Theresa. Speaker at Piksel Festival BE\O/ART, Bergen Norway, Nov. 1 2014 ( “On the Creativity of Primitive Organisms“. -D9.1.4.

Schubert, Theresa. ““, nominated for emerging digital art award, exhibition at Prix Cube Paris, Nov 2015. -D9.3.1.

Schubert, Theresa., & Andrew Adamatzky, Jeff Jones. “Swarmetrical studies: mapping the human body through amorphous intelligence” (Interactive workshop and exhibition, Linz, Sept 2013).-  D.1.3.5.

Schumann, Andrew. “Anti-Platonic Logic and Mathematics.” Journal of Multiple-Valued Logic and Soft Computing 21(1-2) (2013): 53–88. Abstract – D3.3.2.

Schumann, Andrew. “Conventional and Unconventional Reversible Logic Gates on Physarum Polycephalum”, International Journal of Parallel, Emergent and Distributed Systems. -D2.2.8.

Schumann, Andrew. “Game-Theoretic Aspects of Illocutionary Logic, The Eighth Smirnov’s Readings” Proceedings of International Scientific Conference. Moscow, 19–21 June 2013. Moscow, (2013):86–88 – D3.4.3.

Schumann, Andrew. “Non-Linear Permutation Groups on Physarum Polycephalum”, ICSAI 2014, IEEE Xplore, 15‐17 Nov. 2014, pp. 246 – 251. -D7.2.9

Schumann, Andrew. “On Two Squares of Opposition: the Leśniewski’s Style Formalization of Synthetic Propositions” Acta Analytica 28, (2013) 71–93: DOI10.1007/s12136-012-0162-4 On Two Squares of Opposition the Lesniewskis Style Formalization of Synthetic Propositions– D3.1.3.

Schumann, Andrew. “p-Adic Valued Fuzzyness and Experiments with Physarum Polycephalum”, IEEE Xplore, FSKD (2014), p466-472. -D2.2.10

Schumann, Andrew.”Payoff Cellular Automata and Reflexive Games” Journal of Cellular Automata, (2014) Accepted – D3.4.3.

Schumann, Andrew. “Physarum Syllogistic L-Systems”, The Sixth International Conference on Future Computational Technologies and Applications, FUTURE COMPUTING (2014). -D2.2.11.

Schumann, Andrew. “Probabilities on Streams and Reflexive Games, Operations Research and Decisions” (2014) Accepted –  D3.4.2.

Schumann, Andrew . “Proof-Theoretic Cellular Automata as Logic of Unconventional Computing”, International Journal of Unconventional Computing, Vol. 8, No. 3 (2012): 263-280. Abstract

Schumann, Andrew. “Reflexive Games and Non-Archimedean Probabilities, p-Adic Numbers, Ultrametric Analysis and Applications” (2014) Accepted –  D3.4.1.

Schumann, Andrew. “Reversible Logic Gates on Physarum Polycephalum”, Special Volume of the AIP Conference Proceedings, CNAAM 2014, (22‐28 Sep. 2014). -D2.2.12

Schumann, Andrew. “Towards context-based concurrent formal theories”, Parallel Processing Letters. Accepted. -D7.2.2.

Schumann, Andrew.“Towards Slime Mould Based Computer”, New Mathematics and Natural Computation, Accepted. -D7.2.3

Schumann, Andrew. “Unconventional Logics in Decision Making.” Rzeszow: University of Information Technology and Management, January 2014 – D3.1.1.

Schumann, Andrew. “Unconventional logic for massively parallel reasoning, Human System Interaction (HSI)”, The 6th International Conference on 6-8 June 2013, IEEE Xplore, (2013): 298–305 DOI 10.1109/HSI.2013.6577839 – Unconventional Logic for Massively Parrallel Reasoning D3.1.2.

Schumann, A. & Adamatzky, A. “Physarum Polycephalum Diagrams for Syllogistic Systems”, IfCoLog Journal of Logics and their Applications. Accepted . -D2.2.5.

Schumann, A. & Andrew Adamatzky. “The Double-Slit Experiment with Physarum Polycephalum and p-Adic Valued Probabilities and Fuzziness”, International Journal of General Systems. DOI:10.1080/03081079.2014.997530. -D2.2.6.

Schumann, Andrew,  & Andrew Adamatzky.  “Logical Modelling of Physarum Polycephalum”, Analele Universitatii de Vest, Timisoara, Seria Matematica – Informatica XLVIII, 3 (2010): 175-190. arXiv:1105.4060

Schumann,  Andrew,  & Andrew Adamatzky. “Physarum Spatial Logic”, New Math. and Nat. Computation, Vol. 7, No. 3 (2011): 483-498. PDF

Schumann,  Andrew,  & Andrew Adamatzky. “Physarum Polycephalum Diagrams for Syllogistic Systems”, Journal of Logics and their Applications. Accepted. -D9.1.5.

Schumann, Andrew & Pancerz, Krzysztof, Adamatzky, A., Grube, M. “Context-Based Games and Physarum Polycephalum as Simulation Model”, Workshop Unconventional Computation in Europe, Unconventional Computation & Natural Computation 2014, University of Western Ontario, London, Ontario, Canada, July 14-18. -D7.2.6.

Schumann, Andrew & Pancerz, Krzysztof, Adamatzky, A., Grube, M.“Bio-Inspired Game Theory: The Case of Physarum Polycephalum”, BICT 2014, 1‐3 Dec. 2014, ACM. -D7.2.7

Schumann, Andrew, & L Akimova. “Simulating of Schistosomatidae (Trematoda: Digenea) Behaviour by Physarum Spatial Logic.” Annals of Computer Science and Information Systems, Volume 1. Proceedings of the 2013 Federated Conference on Computer Science and Information Systems. IEEE Xplore, (2013):225–2301 – Simulating of Schistosomatidae (Trematoda Digenea) Behavior by Physarum Spatial Logic D3.3.3.

Schumann, A., Akimova, L. “Syllogistic System for the Propagation of Parasites. The Case of Schistosomatidae (Trematoda: Digenea)”, Studies in Logic, Grammar and Rhetoric .-D2.2.7.

Schumann, A., Khrennikov, A. “Quantum Non-Objectivity from Performativity of Quantum Phenomena”, Physica Scripta, Number T163, p15. -D2.2.9

Schumann, Andrew, & Krzysztof Pancerz. “Interfaces in a Game-Theoretic Setting for Controlling the Plasmodium Motions”, BIOSTEC (Biosignals) 2015, Scitepress, 2015, -D7.2.8.

Schumann, Andrew, & Krzysztof Pancerz. “Timed Transition System Models for Programming Physarum Machines: Extended Abstract”, L. Popova-Zeugmann (Ed.), Proceedings of the Workshop on Concurrency, Specification and Programming (CS&P’2014), Chemnitz, Germany, September 29 – October 1, 2014, pp. 180-183. -D7.2.10.

Schumann, Andrew, & Krzysztof Pancerz. “Towards an Object-Oriented Programming Language for Physarum Polycephalum Computing” In: M. Szczuka, L. Czaja, M. Kacprzak (Eds.), Proceedings of the Workshop on Concurrency, Specification and Programming (CS&P’2013), Warsaw, Poland, September 25-27, (2013):389-397 Towards an Object Oriented Programming Language for Physarum Polycephalum Computing – D3.2.1.

Schumann, Andrew, & Krzysztof Pancerz. “Rough Set Approximations of Payoffs in Strategy Games on Physarum Machines. Submitted. -D9.1.8

Schumann, Andrew, & Krzysztof Pancerz. “Interfaces in a Game-Theoretic Setting for Controlling the Plasmodium Motions”, Biosignals (2015). -D9.1.9.

Schumann, Andrew, & Krzysztof Pancerz. “Towards an Object-Oriented Programming Language for Physarum Polycephalum Computing: A Petri Net Model Approach.” Fundamenta Informaticae Submitted – D3.2.2.

Schumann, Andrew, & Krzysztof Pancerz, Jeff Jones. “Towards Logic Circuits Based on Physarum Polycephalum Machines: the Ladder Diagram Approach.” In: A. Cliquet Jr, G. Plantier, T. Schultz, A. Fred, H. Gamboa (Eds.), Proceedings of the 7th International Conference on Biomedical Electronics and Devices (BIODEVICES’2014), Angers, France, March 3-6, (2014) Accepted – D3.3.1.

Schumann, Andrew, & Krzysztof Pancerz, M. Grube, and A. Adamatzky. “Context-Based Games and Physarum Policephalum as Simulation Model”, UCNC (2014). -D9.1.6.

Schumann, A. & K. Pancerz, A. Adamatzky, M. Grube. “Bio-Inspired Game Theory: The Case of Physarum Polycephalum”, BICT (2014). -D9.1.7.

G. Tarabella & P. D’Angelo, A. Cifarelli, A. Dimonte, A. Romeo, T. Berzina, V. Erokhin, and S. Iannotta. “A hybrid living/organic electrochemical transistor based on the Physarum polycephalum cell endowed with both sensing and memristive properties”, Chemical Science (2015). In Press. -D8.2.3.

Tarabella, G., & Angelica Cifarelli, Alice Dimonte, Tatiana Berzina, S. Iannotta, P. D’Angelo. “Physarum Polychephalum with an Electrochemical Cell: the Perspective of a Memristive/Adaptive Response Based on Living Cell Systems.” Proc. E-MRS 2014 Spring meeting, Physica Status solidi C, (2014) Accepted –  D5.1.2

Whiting, James G.H., & Ben De Lacy Costello, Andrew Adamatzky.  “Towards slime mould chemical sensor: Mapping chemical inputs onto electrical potential dynamics of Physarum Polycephalum.” Sensors and Actuators B: Chemical 191, (2014): 844-853  – D1.1.2.



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